Duhan Jasminka Butorac 1. Uvod ................................................................................................................................ 12. Botanička sistematika i podrijetlo duhana ...................................................................... 23. Proizvodnja, površine i prinosi duhana u svijetu i u nas ................................................. 84. Podjela duhana na tipove .............................................................................................. 134.1. Duhani sušeni toplim zrakom u sušnicama (flue-cured) ........................................ 144.2. Duhani sušeni na zraku (air-cured) ........................................................................ 154.3. Duhani sušeni na suncu (sun-cured) ...................................................................... 184.4. Duhani sušeni iznad otvorene vatre (fire-cured) .................................................... 195. Gospodarska važnost duhana ........................................................................................ 206. Morfološka svojstva duhana ......................................................................................... 266.1. Korijen ................................................................................................................... 266.2. Stabljika ................................................................................................................. 266.3. List ......................................................................................................................... 286.4. Cvijet ...................................................................................................................... 296.5. Plod i sjeme ............................................................................................................ 317. Biološka svojstva duhana .............................................................................................. 337.1. Faze razvoja u rasadniku ........................................................................................ 337.1.1. Klijanje sjemena .............................................................................................. 337.1.2. Nicanje ............................................................................................................ 337.1.3. Ukorjenjivanje presadnica .............................................................................. 347.1.4. Formiranje presadnica ..................................................................................... 347.2. Faze razvoja u polju ............................................................................................... 357.2.1. Ukorjenjivanje duhana u polju ........................................................................ 357.2.2. Formiranje biljke ............................................................................................. 357.2.3. Cvjetanje ......................................................................................................... 357.2.4. Formiranje i sazrijevanje sjemena .................................................................. 377.2.5. Formiranje i sazrijevanje listova ..................................................................... 378. Agroekološki uvjeti proizvodnje flue-cured duhana ..................................................... 398.1. Odnos duhana prema temperaturi .......................................................................... 398.2. Odnos duhana prema svjetlu .................................................................................. 408.3. Odnos duhana prema vodi ..................................................................................... 408.4. Odnos duhana prema tlu ........................................................................................ 429. Tehnologija proizvodnje flue-cured duhana ................................................................. 449.1. Uzgoj presadnica duhana klasi?nim na?inom ........................................................ 449.1.1. Priprema lijeha ................................................................................................ 449.1.2. Sjetva ............................................................................................................... 459.1.3. Prihrana ........................................................................................................... 459.1.4. Njega presadnica ............................................................................................. 459.2. Uzgoj presadnica duhana u hidroponima ............................................................... 469.2.1. Priprema hidropona ......................................................................................... 479.2.2. Gnojidba .......................................................................................................... 479.2.3. Sjetva ............................................................................................................... 489.2.4. Njega presadnica ............................................................................................. 509.3. Uzgoj duhana u polju ............................................................................................. 519.3.1. Plodored .......................................................................................................... 519.3.2. Obrada i priprema tla za sadnju ...................................................................... 539.3.3. Gnojidba .......................................................................................................... 549.3.3.1. Dušik ........................................................................................................ 559.3.3.2. Fosfor ....................................................................................................... 589.3.3.3. Kalij .......................................................................................................... 589.3.3.4. Kalcij ........................................................................................................ 599.3.3.5. Magnezij .................................................................................................. 599.3.3.6. Klor .......................................................................................................... 609.3.3.7. Bor ............................................................................................................ 609.3.3.8. Sumpor ..................................................................................................... 609.3.3.9. Mangan .................................................................................................... 619.3.3.10. Ostala mikrohraniva ............................................................................... 619.3.3.11. Unošenje gnojiva ................................................................................... 619.3.4. Ciljevi selekcije i izbor sorte ........................................................................... 629.3.5. Sadnja .............................................................................................................. 699.3.6. Njega duhana .................................................................................................. 729.3.6.1. Suzbijanje korova ..................................................................................... 729.3.6.2. Natapanje ................................................................................................. 749.3.6.3. Otkidanje cvata i zaperaka ....................................................................... 769.3.6.4. Suzbijanje bolesti ..................................................................................... 789.3.6.4.1. Gljivi?ne bolesti ................................................................................ 799.3.6.4.1.1. Polijeganje rasada ili kli?na padavica (Pythium debarianumHesse) ............................................................................................................ 799.3.6.4.1.2. Crna korijenova trulež (Chalara ellegans Naq Raj & W.B.Kendr.) .......................................................................................................... 799.3.6.4.1.3. Plamenja?a (Peronospora tabacina Adam) ............................... 809.3.6.4.1.4. Crna noga (Phytophtora nicotianae (parasitica) Dastur) .......... 809.3.6.4.1.5. Gljivi?no venuüe (Fusarium oxysporum (nicotianae) Schltdl.) 819.3.6.4.1.6. Ranjavost stabljike (Rhizoctonia solani Kuhn) .......................... 819.3.6.4.1.7. Trulež stabljike i korijena duhana (Sclerotium rolfsii Sacc) ...... 819.3.6.4.1.8. Smeya pjegavost (Alternaria alternata (tenuis) Keissl) ............ 819.3.6.4.1.9. Pepelnica (Erisyphe cichoracearum DC) .................................. 829.3.6.4.2. Bakterijske bolesti ............................................................................. 829.3.6.4.2.1. Divlja vatra ili plamac (Pseudomonas tabaci Wolf & Foster) .. 829.3.6.4.2.2. Bakterijsko venuüe (Pseudomonas solanacearum Smith) ......... 839.3.6.4.2.3. Šuplja stabljika (Erwinia carotovora Winslow et al.) ............... 839.3.6.4.3. Virusne bolesti .................................................................................. 839.3.6.4.3.1. Virus mozaika duhana (Tobacco mosaic virus - TMV) ............. 839.3.6.4.3.2. Y virus krumpira (Vein banding - PVY) .................................. 849.3.6.4.3.3. Virus prstenastog venuüa raj?ice (Tomato spotted wilt virus -TSWV) .......................................................................................................... 859.3.6.4.3.4. Virus mozaika krastavca (Cucumber mosaic virus - CMV) ...... 859.3.6.4.3.5. Virusna pjegavost (Tobacco etch virus - TEV) ......................... 869.3.6.4.4. Nematode .......................................................................................... 869.3.6.4.5. Parazitske cvjetnice ........................................................................... 879.3.6.4.5.1. Zumbul ili volovod (Orobanche spp.) ....................................... 879.3.6.4.5.2. Vilina kosica (Cuscuta alba) ..................................................... 889.3.6.4.6. Bolesti fiziološkog podrijetla ............................................................ 889.3.6.5. Suzbijanje štetnika ................................................................................... 889.3.6.5.1. Štetnici u lijehama ............................................................................. 889.3.6.5.1.1. Leteüi duhanski kornjaš ili duhanov buha? (Epitrix hirtipennisMelsh.) .......................................................................................................... 889.3.6.5.1.2. Duhanske mušice (Hydrobaenun spp. Alch.) ............................ 889.3.6.5.1.3. Povrtlarski žižak (Listroderes costriostris obliquus Klug) ........ 899.3.6.5.1.4. Rovac (Gryllotalpa hexadactyla Perty) ..................................... 899.3.6.5.1.5. Zelena lipanjska buba (Cotinis nitida L.) .................................. 899.3.6.5.2. Štetnici u polju .................................................................................. 899.3.6.5.2.1. Duhanski ži?njak (Conoderus vespertinus F. i C. Falli.) ........... 899.3.6.5.2.2. Crna sovica (Agrotis ypsilon Hufnl.) ......................................... 899.3.6.5.2.3. Crv duhanske stabljike (Crambus spp.) ..................................... 899.3.6.5.2.4. Bjeloresa buba (Graphognathus spp.) ....................................... 909.3.6.5.2.5. Duhanski rogati crv (Manduca sexta L.) ................................... 909.3.6.5.2.6. Crv duhanskog pupa (Heliothis virescens F.) ............................ 909.3.6.5.2.7. Breskvina zelena uš (Mysus persicae Sulzer) ............................ 909.3.6.5.2.8. Zrikavac ?etveroto?kasti (Oecanthus nigricornis quadripunctatusBeut) .............................................................................................................. 919.3.6.5.2.9. Crv duhanskog lista ili krumpirov moljac (Phtorimaeaoperculella Zell) ............................................................................................ 919.3.6.5.2.10. Japanski sjajnik (Popilla japonica Newman) .......................... 919.3.6.5.2.11. Kupusni resi?ar (Trichoplusia spp. Hbn.) ................................ 929.3.6.5.2.12. Zelena smrdljiva buba (Nozara viridula L.) ............................ 929.3.6.5.2.13. Duhanov trips resi?ar (Trips tabaci Lind.) .............................. 929.3.6.5.2.14. Mjere suzbijanja štetnika ......................................................... 929.3.6.5.3. Štetnici u skladištu ............................................................................ 949.3.6.5.3.1. Duhanski moljac (Ephestia elutella Hlbn.) ................................ 949.3.6.5.3.2. Duhanski žižak ili duhanar (Lasioderma serricorne F.) ............ 949.3.7. Berba ............................................................................................................... 959.3.8. Sušenje ............................................................................................................ 9810. Dorada duhana .......................................................................................................... 10310.1. Primarna manipulacija ....................................................................................... 10310.2. Otkup duhana ..................................................................................................... 10410.3. Ižiljavanje (thrashing) ........................................................................................ 10610.4. Ponovno sušenje (redrying) ............................................................................... 10810.5. Fermentacija ....................................................................................................... 11010.6. Skladištenje, odležavanje i maturacija ............................................................... 11310.7. Ekspandiranje duhana ........................................................................................ 11410.8. Rekonstituirani duhan (folija) ............................................................................ 11611. Kvaliteta duhana ....................................................................................................... 11811.1. Organolepti?ka procjena .................................................................................... 12111.2. Fizikalna svojstva ............................................................................................... 12711.3. Kemijska svojstva .............................................................................................. 13011.4. Puša?ka svojstva ................................................................................................ 13912. Duhani sušeni na zraku ............................................................................................. 14512.1. Burley ................................................................................................................. 14512.1.1. Agroekološki uvjeti proizvodnje burleya .................................................... 14612.1.2. Tehnologija proizvodnje burleya ................................................................ 14712.2. Maryland ............................................................................................................ 15412.2.1. Agroekološki uvjeti proizvodnje marylanda ............................................... 15512.2.2. Tehnologija proizvodnje marylanda ........................................................... 15512.3. Tamni duhani sušeni na zraku ............................................................................ 15612.3.1. Agroekološki uvjeti proizvodnje tamnih duhana ........................................ 15612.3.2. Tehnologija proizvodnje tamnih duhana .................................................... 15712.4. Cigarni duhani .................................................................................................... 15812.4.1. Agroekološki uvjeti proizvodnje cigarnih duhana ...................................... 15812.4.2. Tehnologija proizvodnje cigarnih duhana .................................................. 15812.5. Perique duhani ................................................................................................... 16112.5.1. Agroekološki uvjeti proizvodnje perique duhana ....................................... 16112.5.2. Tehnologija proizvodnje perique duhana .................................................... 16113. Duhani sušeni na suncu ............................................................................................. 16313.1. Orijentalni duhani .............................................................................................. 16313.1.1. Agroekološki uvjeti proizvodnje orijentalnih duhana ................................. 16313.1.2. Tehnologija proizvodnje orijentalnih duhana ............................................. 16413.2. Poluorijentalni duhani ........................................................................................ 16913.2.1. Agroekološki uvjeti proizvodnje poluorijentalnih duhana ......................... 16913.2.2. Tehnologija proizvodnje poluorijentalnih duhana ...................................... 16913.3. Virginia sun-cured duhani .................................................................................. 17114. Duhani sušeni iznad otvorene vatre .......................................................................... 17214.1. Agroekološki uvjeti proizvodnje fire-cured duhana .......................................... 17214.2. Tehnologija proizvodnje fire-cured duhana ....................................................... 17215. Postupak identifikacije sorata duhana ....................................................................... 17415.1. Morfološka svojstva ........................................................................................... 17415.2. Fenološka svojstva ............................................................................................. 18215.3. Gospodarska svojstva ......................................................................................... 18315.4. Reagiranje na bolesti .......................................................................................... 18415.5. Kemijska svojstva .............................................................................................. 18615.6. Puša?ka svojstva ................................................................................................ 18716. Literatura ................................................................................................................... 189Kazalo pojmova .............................................................................................................. 194